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Having four ridges or crests to the teeth quotations examples
Black bars represent tetralophodont proboscideans; green bars represent dibelodont proboscideans and yellow bar represents tetrabelodont proboscideans (reversion/atavism).
2016 January 13, “The Dance of Tusks: Rediscovery of Lower Incisors in the Pan-American Proboscidean Cuvieronius hyodon Revises Incisor Evolution in Elephantimorpha”, in PLOS ONE